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    Six well known clustering algorithms UPGMA, K-Means, Diana, Fanny, Model-Based and SOM were evaluated.

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    A two-step cluster validation approach is proposed to statistically estimate both the optimal number of clusters and to distinguish significant clusters from noise.

    The resulting clusters reveal coordinated coexpressed genes.

    Most previous studies used function enrichment methods to evaluate modules based on functional annotations, such as GO, MIPS, and KEGG.

    However, some modules may be enriched with too many functions, whereas others may be enriched without any functions, and the background annotation database itself is constantly being updated.

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